首先句子是描述过去一个已经发生的状况,并且对现在有影响。第一个动词用完成时无误。而nearly 99% have vanished 所说的是 也就是these species of living creatures 中的99%遵循与主句时态一致也用完成时,也符合逻辑。
GMAT 语法题目Five hundred million different species of living cr
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