通常文章中的破折号用于表达进一步说明或具体说明,相当于非限定性定语从句 which is/are/means/mean.你的这个句子可用非限定性定语从句取代：
The commonest spaces are those among the particles,which include sand grains and tiny pebbles,of loose,unconsolidated sand and gravel.
英语翻译The commonest spaces are those among the particles—sand
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