Invertebrates (Ray scorpion )had successfully landed before vertebrates .It could have been too late when ichthyostega came onto the land.无脊椎动物先于脊椎动物登陆.在鱼石螈（两栖动物登陆先锋）登陆之前,雷蝎（古节肢动物）已经登陆了.
What were the first animals thought to populate the surface
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