naming,classification,and mapping have also become more particular是主句,因为有谓语have,have之前是主语,后面是宾语
前面那个as引导的原因状语从句,跟and没关系,and只是表示the many thousands of African plant species和their complex ecology的并列关系,在这个原因状语从句中,由于主语the many thousands of African plant species and their complex ecology太长了,为了避免头重脚轻,所以用了倒装句,实际上是As the many thousands of African plant species and their complex ecology more has become known of more这个句子大致意思是,由于好几千种非洲的植物物种和他们复杂的生态学被知道得更多,他们的名字,分类,分布图同样也变得详细了,后面那个则是由动名词引导的从句,表示,不是主句.
希望这么说楼主能明白~>o<
PS.难怪我说后面如此诡异……