当然是有区别的.前者的主语是this ,系动词是is,the museum 是表语.Which引导的部分是修饰the museum这个表语的.后者this museum是主语 ,that 引导的部分是修饰this museum这个主语的,is 是系动词,其他像宾语或表语的句子成分没有出现.不知道这样说你明白了没有.
this is the museum which/in which/at whivh定从.this museum is
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