A pangolin (pronounced /ˈpæŋɡəlɪn/), also scaly anteater or tenggiling, is a mammal of the order Pholidota. There is only one extant family (Manidae) and one genus (Manis) of pangolins, comprising eight species. There are also a number of extinct taxa. Pangolins have large keratin scales covering their skin and are the only mammals with this adaptation. They are found in tropical regions of Africa and Asia. The name "pangolin" derives from the Malay word pengguling ("something that rolls up"). Pangolins are nocturnal animals, and use their well-developed sense of smell to find insects. The long-tailed pangolin is also active by day. Pangolins spend most of the daytime sleeping, curled up into a ball.
The physical appearance of pangolins is marked by large, hardened, plate-like scales. The scales, which are soft on newborn pangolins but harden as the animal matures, are made of keratin, the same material of which human fingernails and tetrapod claws are made. The pangolin is often compared to a walking pine cone or globe artichoke. It can curl up into a ball when threatened, with its overlapping scales acting as armour and its face tucked under its tail. The scales are razor-sharp, providing extra defence. The front claws are so long that they are unsuited for walking, and so the animal walks with its fore paws curled over to protect them. Pangolins can also emit a noxious smelling acid from glands near the anus, similar to the spray of a skunk. Pangolins have short legs, with sharp claws which they use for burrowing into termite and ant mounds, as well as climbing.
The size of pangolins varies by species, ranging from 30 cm to 100 cm (12 to 39 inches). Females are generally smaller than males.
The tongues of pangolins are extremely elongated and extend into the abdominal cavity. By convergent evolution pangolins, the giant anteater, and the tube-lipped nectar bat, all have tongues which are unattached from their hyoid bone and extend past their pharynx deep into the thorax. This extension lies between the sternum and the trachea. Large pangolins can extend their tongues as much as 40 cm (16 inches), with a diameter of only 0.5 cm (1/4 inch).
In pangolins, the section of the brain that relates to problem solving is highly developed. Although their problem solving ability is primarily used to find food in obscure locations, when kept in captivity pangolins are remarkable escape artists.
Arboreal pangolins live in hollow trees, whereas the ground dwelling species dig tunnels underground, up to a depth of 3.5 m (11 feet). Pangolins are also good swimmers.